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 <front>
  <journal-meta>
   <journal-id journal-id-type="publisher-id">Russian Journal of Biological Physics and Chemisrty</journal-id>
   <journal-title-group>
    <journal-title xml:lang="en">Russian Journal of Biological Physics and Chemisrty</journal-title>
    <trans-title-group xml:lang="ru">
     <trans-title>АКТУАЛЬНЫЕ ВОПРОСЫ БИОЛОГИЧЕСКОЙ ФИЗИКИ И ХИМИИ</trans-title>
    </trans-title-group>
   </journal-title-group>
   <issn publication-format="print">2499-9962</issn>
  </journal-meta>
  <article-meta>
   <article-id pub-id-type="publisher-id">54710</article-id>
   <article-categories>
    <subj-group subj-group-type="toc-heading" xml:lang="ru">
     <subject>ЭКОЛОГИЧЕСКАЯ БИОФИЗИКА</subject>
    </subj-group>
    <subj-group subj-group-type="toc-heading" xml:lang="en">
     <subject>ECOLOGICAL BIOPHYSICS</subject>
    </subj-group>
    <subj-group>
     <subject>ЭКОЛОГИЧЕСКАЯ БИОФИЗИКА</subject>
    </subj-group>
   </article-categories>
   <title-group>
    <article-title xml:lang="en">Ecological biophysics - possible horizons of development</article-title>
    <trans-title-group xml:lang="ru">
     <trans-title>Экологическая биофизика - возможные горизонты развития</trans-title>
    </trans-title-group>
   </title-group>
   <contrib-group content-type="authors">
    <contrib contrib-type="author">
     <name-alternatives>
      <name xml:lang="ru">
       <surname>Барцев</surname>
       <given-names>С И</given-names>
      </name>
      <name xml:lang="en">
       <surname>Bartsev</surname>
       <given-names>S I</given-names>
      </name>
     </name-alternatives>
     <email>bartsevsi@ibp.ru</email>
     <xref ref-type="aff" rid="aff-1"/>
    </contrib>
    <contrib contrib-type="author">
     <name-alternatives>
      <name xml:lang="ru">
       <surname>Дегерменджи</surname>
       <given-names>А Г</given-names>
      </name>
      <name xml:lang="en">
       <surname>Degermendzhi</surname>
       <given-names>A G</given-names>
      </name>
     </name-alternatives>
     <xref ref-type="aff" rid="aff-2"/>
    </contrib>
    <contrib contrib-type="author">
     <name-alternatives>
      <name xml:lang="ru">
       <surname>Дегерменджи</surname>
       <given-names>Н Н</given-names>
      </name>
      <name xml:lang="en">
       <surname>Degermendzhi</surname>
       <given-names>N N</given-names>
      </name>
     </name-alternatives>
     <xref ref-type="aff" rid="aff-3"/>
    </contrib>
    <contrib contrib-type="author">
     <name-alternatives>
      <name xml:lang="ru">
       <surname>Салтыков</surname>
       <given-names>М Ю</given-names>
      </name>
      <name xml:lang="en">
       <surname>Saltykov</surname>
       <given-names>M Yu</given-names>
      </name>
     </name-alternatives>
     <xref ref-type="aff" rid="aff-4"/>
    </contrib>
    <contrib contrib-type="author">
     <name-alternatives>
      <name xml:lang="ru">
       <surname>Сарангова</surname>
       <given-names>А Б</given-names>
      </name>
      <name xml:lang="en">
       <surname>Sarangova</surname>
       <given-names>A B</given-names>
      </name>
     </name-alternatives>
     <xref ref-type="aff" rid="aff-5"/>
    </contrib>
   </contrib-group>
   <aff-alternatives id="aff-1">
    <aff>
     <institution xml:lang="ru">Институт биофизики СО РАН, ФИЦ КНЦ СО РАН; Сибирский федеральный университет</institution>
     <country>ru</country>
    </aff>
    <aff>
     <institution xml:lang="en">Institute of Biophysics SB RAS, Federal Research Center KSC SB RAS; Siberian Federal University</institution>
     <country>ru</country>
    </aff>
   </aff-alternatives>
   <aff-alternatives id="aff-2">
    <aff>
     <institution xml:lang="ru">Институт биофизики СО РАН, ФИЦ КНЦ СО РАН</institution>
     <country>ru</country>
    </aff>
    <aff>
     <institution xml:lang="en">Institute of Biophysics SB RAS, Federal Research Center KSC SB RAS</institution>
     <country>ru</country>
    </aff>
   </aff-alternatives>
   <aff-alternatives id="aff-3">
    <aff>
     <institution xml:lang="ru">Красноярский государственный медицинский университет</institution>
     <country>ru</country>
    </aff>
    <aff>
     <institution xml:lang="en">Krasnoyarsk State Medical University</institution>
     <country>ru</country>
    </aff>
   </aff-alternatives>
   <aff-alternatives id="aff-4">
    <aff>
     <institution xml:lang="ru">Институт биофизики СО РАН, ФИЦ КНЦ СО РАН; Сибирский федеральный университет</institution>
     <country>ru</country>
    </aff>
    <aff>
     <institution xml:lang="en">Institute of Biophysics SB RAS, Federal Research Center KSC SB RAS; Siberian Federal University</institution>
     <country>ru</country>
    </aff>
   </aff-alternatives>
   <aff-alternatives id="aff-5">
    <aff>
     <institution xml:lang="ru">Сибирский федеральный университет</institution>
     <country>ru</country>
    </aff>
    <aff>
     <institution xml:lang="en">Siberian Federal University</institution>
     <country>ru</country>
    </aff>
   </aff-alternatives>
   <pub-date publication-format="print" date-type="pub" iso-8601-date="2021-12-25T20:22:29+03:00">
    <day>25</day>
    <month>12</month>
    <year>2021</year>
   </pub-date>
   <pub-date publication-format="electronic" date-type="pub" iso-8601-date="2021-12-25T20:22:29+03:00">
    <day>25</day>
    <month>12</month>
    <year>2021</year>
   </pub-date>
   <volume>6</volume>
   <issue>4</issue>
   <fpage>665</fpage>
   <lpage>674</lpage>
   <history>
    <date date-type="received" iso-8601-date="2021-12-20T20:22:29+03:00">
     <day>20</day>
     <month>12</month>
     <year>2021</year>
    </date>
    <date date-type="accepted" iso-8601-date="2021-12-20T20:22:29+03:00">
     <day>20</day>
     <month>12</month>
     <year>2021</year>
    </date>
   </history>
   <self-uri xlink:href="https://rusjbpc.ru/en/nauka/article/54710/view">https://rusjbpc.ru/en/nauka/article/54710/view</self-uri>
   <abstract xml:lang="ru">
    <p>В статье обсуждается специфика подхода экологической биофизики к исследования экологических систем и земной биосферы в целом. Рассмотрены характерные для этого раздела биофизики проблемы и возможные подходы к их преодолению. В частности, проблема уникальности экологических систем может быть существенно ослаблена, если удастся разработать теорию экологического подобия. Проблема разнонаправленной зависимости устойчивости реальных экосистем и их моделей от количества входящих видов может быть снята путем перехода к моделям, так называемого, гибкого метаболизма. Необозримая сложность биосферы побуждает обращаться к концептуальным моделям, построенным по принципу наихудшего сценария, которые позволяют, хотя бы в общем виде, оценить действие и возможный вклад тех или иных механизмов в глобальную динамику.</p>
   </abstract>
   <trans-abstract xml:lang="en">
    <p>The paper discusses the specificity of the approach of ecological biophysics to the study of ecological systems and the Earth's biosphere in general. The problems typical for this section of biophysics and possible approaches to their overcoming are considered. In particular, the problem of the uniqueness of ecological systems can be significantly weakened if it is possible to develop a theory of ecological similarity. The problem of the multidirectional dependence of the stability of real ecosystems and their models on the number of incoming species can be removed by switching to the models of the so-called flexible metabolism. The immense complexity of the biosphere prompts one to turn to conceptual models based on the principle of the worst-case scenario, which allow, at least in general terms, to assess the action and possible contribution of certain mechanisms to global dynamics.</p>
   </trans-abstract>
   <kwd-group xml:lang="ru">
    <kwd>экологическая биофизика</kwd>
    <kwd>экосистема</kwd>
    <kwd>аутостабилизация</kwd>
    <kwd>биосфера</kwd>
   </kwd-group>
   <kwd-group xml:lang="en">
    <kwd>ecological biophysics</kwd>
    <kwd>ecosystem</kwd>
    <kwd>autostabilization</kwd>
    <kwd>biosphere</kwd>
   </kwd-group>
  </article-meta>
 </front>
 <body>
  <p></p>
 </body>
 <back>
  <ref-list>
   <ref id="B1">
    <label>1.</label>
    <citation-alternatives>
     <mixed-citation xml:lang="ru">
            
              Моровиц Г. Исторический очерк. Теоретическая и математическая биология. М.: Мир, 1968, с. 34-48. [Morowitz G. Historical sketch. Theoretical and mathematical biology. M.: Mir, 1968, pp. 34-48. (In Russ.)]
            
          </mixed-citation>
     <mixed-citation xml:lang="en">
            
              Morovic G. Istoricheskiy ocherk. Teoreticheskaya i matematicheskaya biologiya. M.: Mir, 1968, s. 34-48. [Morowitz G. Historical sketch. Theoretical and mathematical biology. M.: Mir, 1968, pp. 34-48. (In Russ.)]
            
          </mixed-citation>
    </citation-alternatives>
   </ref>
   <ref id="B2">
    <label>2.</label>
    <citation-alternatives>
     <mixed-citation xml:lang="ru">
            
              Вернадский В.И. Биосфера и ноосфера. Айрис-пресс, 2012, 576 с. [Vernadsky V.I. Biosphere and noosphere. Iris press, 2012, 576 p. (In Russ.)]
            
          </mixed-citation>
     <mixed-citation xml:lang="en">
            
              Vernadskiy V.I. Biosfera i noosfera. Ayris-press, 2012, 576 s. [Vernadsky V.I. Biosphere and noosphere. Iris press, 2012, 576 p. (In Russ.)]
            
          </mixed-citation>
    </citation-alternatives>
   </ref>
   <ref id="B3">
    <label>3.</label>
    <citation-alternatives>
     <mixed-citation xml:lang="ru">
            
              Degermendzhi A.G. New directions in biophysical ecology. Chapter 14 Ch 10. In: Global Climatology and Ecodynamics: Anthropogenic Changes to Planet Earth (Springer Praxis Books/Environmental Sciences), 2009, pp. 379-396.
            
          </mixed-citation>
     <mixed-citation xml:lang="en">
            
              Degermendzhi A.G. New directions in biophysical ecology. Chapter 14 Ch 10. In: Global Climatology and Ecodynamics: Anthropogenic Changes to Planet Earth (Springer Praxis Books/Environmental Sciences), 2009, pp. 379-396.
            
          </mixed-citation>
    </citation-alternatives>
   </ref>
   <ref id="B4">
    <label>4.</label>
    <citation-alternatives>
     <mixed-citation xml:lang="ru">
            
              Барцев С.И., Барцева О.Д. Симметрии структуры и эквифинальность эволюционных исходов в простых нейросетевых моделях. ДАН, 2002, т. 386, № 1, с. 114-117. [Bartsev S.I., Bartseva O.D.Structure symmetries and equifinality of evolutionary outcomes in simple neural network models. DAN, 2002, vol. 386, no. 1, pp. 114-117. (In Russ.)]
            
          </mixed-citation>
     <mixed-citation xml:lang="en">
            
              Barcev S.I., Barceva O.D. Simmetrii struktury i ekvifinal'nost' evolyucionnyh ishodov v prostyh neyrosetevyh modelyah. DAN, 2002, t. 386, № 1, s. 114-117. [Bartsev S.I., Bartseva O.D.Structure symmetries and equifinality of evolutionary outcomes in simple neural network models. DAN, 2002, vol. 386, no. 1, pp. 114-117. (In Russ.)]
            
          </mixed-citation>
    </citation-alternatives>
   </ref>
   <ref id="B5">
    <label>5.</label>
    <citation-alternatives>
     <mixed-citation xml:lang="ru">
            
              Барцев С.И., Барцева О.Д. Функционально-инвариантный подход к проблеме уникальности биологических систем: простая нейросетевая модель. ДАН, 2005, т. 405, № 4, с. 1-4. [Bartsev S.I., Bartseva O.D. Functionally invariant approach to the problem of the uniqueness of biological systems: a simple neural network model. DAN, 2005, vol. 405, no. 4, pp. 1-4. (In Russ.)]
            
          </mixed-citation>
     <mixed-citation xml:lang="en">
            
              Barcev S.I., Barceva O.D. Funkcional'no-invariantnyy podhod k probleme unikal'nosti biologicheskih sistem: prostaya neyrosetevaya model'. DAN, 2005, t. 405, № 4, s. 1-4. [Bartsev S.I., Bartseva O.D. Functionally invariant approach to the problem of the uniqueness of biological systems: a simple neural network model. DAN, 2005, vol. 405, no. 4, pp. 1-4. (In Russ.)]
            
          </mixed-citation>
    </citation-alternatives>
   </ref>
   <ref id="B6">
    <label>6.</label>
    <citation-alternatives>
     <mixed-citation xml:lang="ru">
            
              Барцев С.И., Барцева О.Д. Эвристические нейросетевые моделив биофизике: приложение к проблеме структурно-функционального соответствия. Красноярск: Сибирский федеральный ун-т, 2010, 115 с. [Bartsev S.I., Bartseva O.D. Heuristic neural network models in biophysics: application to the problem of structural and functional correspondence. Krasnoyarsk: Siberian Federal University, 2010, 115 p. (In Russ.)]
            
          </mixed-citation>
     <mixed-citation xml:lang="en">
            
              Barcev S.I., Barceva O.D. Evristicheskie neyrosetevye modeliv biofizike: prilozhenie k probleme strukturno-funkcional'nogo sootvetstviya. Krasnoyarsk: Sibirskiy federal'nyy un-t, 2010, 115 s. [Bartsev S.I., Bartseva O.D. Heuristic neural network models in biophysics: application to the problem of structural and functional correspondence. Krasnoyarsk: Siberian Federal University, 2010, 115 p. (In Russ.)]
            
          </mixed-citation>
    </citation-alternatives>
   </ref>
   <ref id="B7">
    <label>7.</label>
    <citation-alternatives>
     <mixed-citation xml:lang="ru">
            
              Свирежев Ю.М. Нелинейные волны, диссипативные структуры и катастрофы в экологии. М.: Наука, 1987, 386 с. [Svirezhev Yu.M. Nonlinear waves, dissipative structures and catastrophes in ecology. Moscow: Nauka, 1987, 386 p. (In Russ.)]
            
          </mixed-citation>
     <mixed-citation xml:lang="en">
            
              Svirezhev Yu.M. Nelineynye volny, dissipativnye struktury i katastrofy v ekologii. M.: Nauka, 1987, 386 s. [Svirezhev Yu.M. Nonlinear waves, dissipative structures and catastrophes in ecology. Moscow: Nauka, 1987, 386 p. (In Russ.)]
            
          </mixed-citation>
    </citation-alternatives>
   </ref>
   <ref id="B8">
    <label>8.</label>
    <citation-alternatives>
     <mixed-citation xml:lang="ru">
            
              Адамович В.В., Дегерменджи А.Г. Статистические закономерности организации маловидовых стационарных сообществ микроорганизмов. Ж. Общей биологии, 1985, т. XLVI, № 4, c. 527-532. [Adamovich V.V., Degermendzhi A.G. Statistical regularities of the organization of small-species stationary communities of microorganisms. J. General Biology, 1985, vol. XLVI, no. 4, pp. 527-532. (In Russ.)]
            
          </mixed-citation>
     <mixed-citation xml:lang="en">
            
              Adamovich V.V., Degermendzhi A.G. Statisticheskie zakonomernosti organizacii malovidovyh stacionarnyh soobschestv mikroorganizmov. Zh. Obschey biologii, 1985, t. XLVI, № 4, c. 527-532. [Adamovich V.V., Degermendzhi A.G. Statistical regularities of the organization of small-species stationary communities of microorganisms. J. General Biology, 1985, vol. XLVI, no. 4, pp. 527-532. (In Russ.)]
            
          </mixed-citation>
    </citation-alternatives>
   </ref>
   <ref id="B9">
    <label>9.</label>
    <citation-alternatives>
     <mixed-citation xml:lang="ru">
            
              Дегерменджи А.Г., Абакумов А.И. Принцип конкурентного исключения в двухвидовом сообществе с одним метаболическим фактором регуляции. ДАН, 2018, т. 480, № 4, с. 495-498. [Degermendzhi A.G., Abakumov A.I. The principle of competitive exclusion in a two-species community with one metabolic regulation factor. DAN, 2018, vol. 480, no. 4, pp. 495-498. (In Russ.)]
            
          </mixed-citation>
     <mixed-citation xml:lang="en">
            
              Degermendzhi A.G., Abakumov A.I. Princip konkurentnogo isklyucheniya v dvuhvidovom soobschestve s odnim metabolicheskim faktorom regulyacii. DAN, 2018, t. 480, № 4, s. 495-498. [Degermendzhi A.G., Abakumov A.I. The principle of competitive exclusion in a two-species community with one metabolic regulation factor. DAN, 2018, vol. 480, no. 4, pp. 495-498. (In Russ.)]
            
          </mixed-citation>
    </citation-alternatives>
   </ref>
   <ref id="B10">
    <label>10.</label>
    <citation-alternatives>
     <mixed-citation xml:lang="ru">
            
              Дегерменджи А.Г., Печуркин Н.С., Шкидченко А.Н. Аутостабилизация факторов, контролирующих рост в биологических системах. Новосибирск: Наука, 1979, 141 с. [Degermendzhi A.G., Pechurkin N.S., Shkidchenko A.N. Autostabilization of factors that control growth in biological systems. Novosibirsk: Science, 1979, 141 p. (In Russ.)]
            
          </mixed-citation>
     <mixed-citation xml:lang="en">
            
              Degermendzhi A.G., Pechurkin N.S., Shkidchenko A.N. Autostabilizaciya faktorov, kontroliruyuschih rost v biologicheskih sistemah. Novosibirsk: Nauka, 1979, 141 s. [Degermendzhi A.G., Pechurkin N.S., Shkidchenko A.N. Autostabilization of factors that control growth in biological systems. Novosibirsk: Science, 1979, 141 p. (In Russ.)]
            
          </mixed-citation>
    </citation-alternatives>
   </ref>
   <ref id="B11">
    <label>11.</label>
    <citation-alternatives>
     <mixed-citation xml:lang="ru">
            
              Адамович В.А., Терсков И.А., Дегерменджи А.Г. Эффект ауто стабилизации контролирующих рост факторов и взаимодействия в сообществе. ДАН, 1987, т. 235, № 5, с. 1236-1239. [Adamovich V.A., Terskov I.A., Degermendzhi A.G. The effect of self-stabilizing growth control factors and interactions in the community. DAN, 1987, vol. 235, no. 5, pp. 1236-1239. (In Russ.)]
            
          </mixed-citation>
     <mixed-citation xml:lang="en">
            
              Adamovich V.A., Terskov I.A., Degermendzhi A.G. Effekt auto stabilizacii kontroliruyuschih rost faktorov i vzaimodeystviya v soobschestve. DAN, 1987, t. 235, № 5, s. 1236-1239. [Adamovich V.A., Terskov I.A., Degermendzhi A.G. The effect of self-stabilizing growth control factors and interactions in the community. DAN, 1987, vol. 235, no. 5, pp. 1236-1239. (In Russ.)]
            
          </mixed-citation>
    </citation-alternatives>
   </ref>
   <ref id="B12">
    <label>12.</label>
    <citation-alternatives>
     <mixed-citation xml:lang="ru">
            
              Hutchinson G.E. The paradox of the plankton. The American Naturalist, 1961, vol. 95, no. 882, pp. 137-145.
            
          </mixed-citation>
     <mixed-citation xml:lang="en">
            
              Hutchinson G.E. The paradox of the plankton. The American Naturalist, 1961, vol. 95, no. 882, pp. 137-145.
            
          </mixed-citation>
    </citation-alternatives>
   </ref>
   <ref id="B13">
    <label>13.</label>
    <citation-alternatives>
     <mixed-citation xml:lang="ru">
            
              Levine J.M., HilleRisLambers J. The importance of niches for the maintenance of species diversity. Nature, 2009, vol. 461, pp. 254-257.
            
          </mixed-citation>
     <mixed-citation xml:lang="en">
            
              Levine J.M., HilleRisLambers J. The importance of niches for the maintenance of species diversity. Nature, 2009, vol. 461, pp. 254-257.
            
          </mixed-citation>
    </citation-alternatives>
   </ref>
   <ref id="B14">
    <label>14.</label>
    <citation-alternatives>
     <mixed-citation xml:lang="ru">
            
              Свирежев Ю.М., Логофет Д.О. Устойчивость биологических сообществ. М.: Наука, 1978, 352 с. [Svirezhev Yu.M., Logofet D.O. Sustainability of biological communities. M.: Nauka, 1978, 352 p. (In Russ.)]
            
          </mixed-citation>
     <mixed-citation xml:lang="en">
            
              Svirezhev Yu.M., Logofet D.O. Ustoychivost' biologicheskih soobschestv. M.: Nauka, 1978, 352 s. [Svirezhev Yu.M., Logofet D.O. Sustainability of biological communities. M.: Nauka, 1978, 352 p. (In Russ.)]
            
          </mixed-citation>
    </citation-alternatives>
   </ref>
   <ref id="B15">
    <label>15.</label>
    <citation-alternatives>
     <mixed-citation xml:lang="ru">
            
              Winemiller K.O. Spatial and temporal variation in tropical fish trophic networks. Ecol.Monogr., 1990, vol. 60, pp. 331-367.
            
          </mixed-citation>
     <mixed-citation xml:lang="en">
            
              Winemiller K.O. Spatial and temporal variation in tropical fish trophic networks. Ecol.Monogr., 1990, vol. 60, pp. 331-367.
            
          </mixed-citation>
    </citation-alternatives>
   </ref>
   <ref id="B16">
    <label>16.</label>
    <citation-alternatives>
     <mixed-citation xml:lang="ru">
            
              Polis G.A.Complex trophic interactions in deserts: an empirical critique of food web theory. The American Naturalist, 1991, vol. 138, pp. 123-155.
            
          </mixed-citation>
     <mixed-citation xml:lang="en">
            
              Polis G.A.Complex trophic interactions in deserts: an empirical critique of food web theory. The American Naturalist, 1991, vol. 138, pp. 123-155.
            
          </mixed-citation>
    </citation-alternatives>
   </ref>
   <ref id="B17">
    <label>17.</label>
    <citation-alternatives>
     <mixed-citation xml:lang="ru">
            
              Ives A.R., Carpenter S.R. Stability and diversity of ecosystems. Science, 2007, vol. 317, pp. 58-62.
            
          </mixed-citation>
     <mixed-citation xml:lang="en">
            
              Ives A.R., Carpenter S.R. Stability and diversity of ecosystems. Science, 2007, vol. 317, pp. 58-62.
            
          </mixed-citation>
    </citation-alternatives>
   </ref>
   <ref id="B18">
    <label>18.</label>
    <citation-alternatives>
     <mixed-citation xml:lang="ru">
            
              Bartsev S.I. Stoichiometric constraints and complete closure of long-term life support systems. Adv Space Res., 2004, vol. 34, no. 7, pp. 1509-1516.
            
          </mixed-citation>
     <mixed-citation xml:lang="en">
            
              Bartsev S.I. Stoichiometric constraints and complete closure of long-term life support systems. Adv Space Res., 2004, vol. 34, no. 7, pp. 1509-1516.
            
          </mixed-citation>
    </citation-alternatives>
   </ref>
   <ref id="B19">
    <label>19.</label>
    <citation-alternatives>
     <mixed-citation xml:lang="ru">
            
              Салтыков М.Ю., Барцев С.И., Ланкин Ю.П. Зависимость устойчивости моделей замкнутых экосистем от числа видов. Журнал СФУ серия «Биология 2», 2011, № 4, с. 197-208. [Saltykov M.Yu., Bartsev S.I., Lankin Yu.P. Dependence of the stability of models of closed ecosystems on the number of species. SibFU journal series &quot;Biology 2&quot;, 2011, no. 4, pp. 197-208. (In Russ.)]
            
          </mixed-citation>
     <mixed-citation xml:lang="en">
            
              Saltykov M.Yu., Barcev S.I., Lankin Yu.P. Zavisimost' ustoychivosti modeley zamknutyh ekosistem ot chisla vidov. Zhurnal SFU seriya «Biologiya 2», 2011, № 4, s. 197-208. [Saltykov M.Yu., Bartsev S.I., Lankin Yu.P. Dependence of the stability of models of closed ecosystems on the number of species. SibFU journal series &quot;Biology 2&quot;, 2011, no. 4, pp. 197-208. (In Russ.)]
            
          </mixed-citation>
    </citation-alternatives>
   </ref>
   <ref id="B20">
    <label>20.</label>
    <citation-alternatives>
     <mixed-citation xml:lang="ru">
            
              Saltykov M.Yu., Bartsev S.I., Lankin Yu.P. Stability of CELSS models as dependent upon the properties of methabolism of the described species. Adv. Space Res., 2012, vol. 49, no. 2, pp. 229-223.
            
          </mixed-citation>
     <mixed-citation xml:lang="en">
            
              Saltykov M.Yu., Bartsev S.I., Lankin Yu.P. Stability of CELSS models as dependent upon the properties of methabolism of the described species. Adv. Space Res., 2012, vol. 49, no. 2, pp. 229-223.
            
          </mixed-citation>
    </citation-alternatives>
   </ref>
   <ref id="B21">
    <label>21.</label>
    <citation-alternatives>
     <mixed-citation xml:lang="ru">
            
              Saltykov M.Yu., Bartsev S.I. Developing of discrimination experiment to find most adequate model of plant’s multi-nutrient functional response. IOP Conf. Series: Materials Science and Engineering, 2017, vol. 173, p. 012017. doi: 10.1088/1757-899X/173/1/012017
            
          </mixed-citation>
     <mixed-citation xml:lang="en">
            
              Saltykov M.Yu., Bartsev S.I. Developing of discrimination experiment to find most adequate model of plant’s multi-nutrient functional response. IOP Conf. Series: Materials Science and Engineering, 2017, vol. 173, p. 012017. doi: 10.1088/1757-899X/173/1/012017
            
          </mixed-citation>
    </citation-alternatives>
   </ref>
   <ref id="B22">
    <label>22.</label>
    <citation-alternatives>
     <mixed-citation xml:lang="ru">
            
              Bartsev S.I., Degermendzhi A.G., Sarangova A.B. Closure of Earth’s Biosphere: Evolution and Current State. Journal of Siberian Federal University. Biology, 2019 vol. 12, no. 3, pp. 337-347.
            
          </mixed-citation>
     <mixed-citation xml:lang="en">
            
              Bartsev S.I., Degermendzhi A.G., Sarangova A.B. Closure of Earth’s Biosphere: Evolution and Current State. Journal of Siberian Federal University. Biology, 2019 vol. 12, no. 3, pp. 337-347.
            
          </mixed-citation>
    </citation-alternatives>
   </ref>
   <ref id="B23">
    <label>23.</label>
    <citation-alternatives>
     <mixed-citation xml:lang="ru">
            
              Marcott S.A., Shakun J.D., Clark P.U., Mix A.C. A Reconstruction of Regional and Global Temperature for the Past 11,300 Years. Science, 2013, vol. 339, pp. 1198-1201.
            
          </mixed-citation>
     <mixed-citation xml:lang="en">
            
              Marcott S.A., Shakun J.D., Clark P.U., Mix A.C. A Reconstruction of Regional and Global Temperature for the Past 11,300 Years. Science, 2013, vol. 339, pp. 1198-1201.
            
          </mixed-citation>
    </citation-alternatives>
   </ref>
   <ref id="B24">
    <label>24.</label>
    <citation-alternatives>
     <mixed-citation xml:lang="ru">
            
              Neukom R., Steiger N., Gómez-Navarro J.J., Wang J., Werner J.P. No evidence for globally coherent warm and cold periods over the preindustrial Common Era. Nature, 2019, vol. 571, pp.550-572.
            
          </mixed-citation>
     <mixed-citation xml:lang="en">
            
              Neukom R., Steiger N., Gómez-Navarro J.J., Wang J., Werner J.P. No evidence for globally coherent warm and cold periods over the preindustrial Common Era. Nature, 2019, vol. 571, pp.550-572.
            
          </mixed-citation>
    </citation-alternatives>
   </ref>
   <ref id="B25">
    <label>25.</label>
    <citation-alternatives>
     <mixed-citation xml:lang="ru">
            
              Drijfhout S. et al. Catalogue of abrupt shifts in Intergovernmental Panel on Climate Change climate models. PNAS, 2015, vol. 112, no. 43, pp. E5777-E5786.
            
          </mixed-citation>
     <mixed-citation xml:lang="en">
            
              Drijfhout S. et al. Catalogue of abrupt shifts in Intergovernmental Panel on Climate Change climate models. PNAS, 2015, vol. 112, no. 43, pp. E5777-E5786.
            
          </mixed-citation>
    </citation-alternatives>
   </ref>
   <ref id="B26">
    <label>26.</label>
    <citation-alternatives>
     <mixed-citation xml:lang="ru">
            
              Lenton T.M. et al. Climate tipping points-Too risky to bet against. Nature, 2019, vol. 575, pp. 592-595.
            
          </mixed-citation>
     <mixed-citation xml:lang="en">
            
              Lenton T.M. et al. Climate tipping points-Too risky to bet against. Nature, 2019, vol. 575, pp. 592-595.
            
          </mixed-citation>
    </citation-alternatives>
   </ref>
   <ref id="B27">
    <label>27.</label>
    <citation-alternatives>
     <mixed-citation xml:lang="ru">
            
              Steffen W. et al. Trajectories of the Earth System in the Anthropocene. PNAS, 2018, vol. 115, no. 33, pp. 8252-8259.
            
          </mixed-citation>
     <mixed-citation xml:lang="en">
            
              Steffen W. et al. Trajectories of the Earth System in the Anthropocene. PNAS, 2018, vol. 115, no. 33, pp. 8252-8259.
            
          </mixed-citation>
    </citation-alternatives>
   </ref>
   <ref id="B28">
    <label>28.</label>
    <citation-alternatives>
     <mixed-citation xml:lang="ru">
            
              Барцев С.И., Дегерменджи А.Г. Ерохин Д.В. Глобальная минимальная модель многолетней динамики углерода в биосфере. ДАН, 2005, т. 401, № 2, с. 233-237. [Bartsev S.I., Degermendzhi A.G., Erokhin D.V. Global minimum model of long-term dynamics of carbon in the biosphere. DAN, 2005, vol. 401, no. 2, pp. 233-237. (In Russ.)]
            
          </mixed-citation>
     <mixed-citation xml:lang="en">
            
              Barcev S.I., Degermendzhi A.G. Erohin D.V. Global'naya minimal'naya model' mnogoletney dinamiki ugleroda v biosfere. DAN, 2005, t. 401, № 2, s. 233-237. [Bartsev S.I., Degermendzhi A.G., Erokhin D.V. Global minimum model of long-term dynamics of carbon in the biosphere. DAN, 2005, vol. 401, no. 2, pp. 233-237. (In Russ.)]
            
          </mixed-citation>
    </citation-alternatives>
   </ref>
   <ref id="B29">
    <label>29.</label>
    <citation-alternatives>
     <mixed-citation xml:lang="ru">
            
              Bartsev S.I., Degermendzhi A.G., Erokhin D.V. Principle of the worst scenario in the modelling past and future of biosphere dynamics. Ecological Modelling, 2008, vol. 216, no. 2, pp. 160-171.
            
          </mixed-citation>
     <mixed-citation xml:lang="en">
            
              Bartsev S.I., Degermendzhi A.G., Erokhin D.V. Principle of the worst scenario in the modelling past and future of biosphere dynamics. Ecological Modelling, 2008, vol. 216, no. 2, pp. 160-171.
            
          </mixed-citation>
    </citation-alternatives>
   </ref>
   <ref id="B30">
    <label>30.</label>
    <citation-alternatives>
     <mixed-citation xml:lang="ru">
            
              Кондратьев К.Я., Крапивин В.Ф. Моделирование глобального круговорота углерода. М.: Физматлит, 2004, 336 с. [Kondratyev K.Ya., Krapivin V.F. Modeling the global carbon cycle. Moscow: Fizmatlit, 2004, 336 p. (In Russ.)]
            
          </mixed-citation>
     <mixed-citation xml:lang="en">
            
              Kondrat'ev K.Ya., Krapivin V.F. Modelirovanie global'nogo krugovorota ugleroda. M.: Fizmatlit, 2004, 336 s. [Kondratyev K.Ya., Krapivin V.F. Modeling the global carbon cycle. Moscow: Fizmatlit, 2004, 336 p. (In Russ.)]
            
          </mixed-citation>
    </citation-alternatives>
   </ref>
   <ref id="B31">
    <label>31.</label>
    <citation-alternatives>
     <mixed-citation xml:lang="ru">
            
              IPCC, 2001. Climate Change. 2001: Scientific aspects, UNEP, 881 p.
            
          </mixed-citation>
     <mixed-citation xml:lang="en">
            
              IPCC, 2001. Climate Change. 2001: Scientific aspects, UNEP, 881 p.
            
          </mixed-citation>
    </citation-alternatives>
   </ref>
   <ref id="B32">
    <label>32.</label>
    <citation-alternatives>
     <mixed-citation xml:lang="ru">
            
              Brovkin V., Sitch S., Bloh von W., Claussen M., Bauer E., Cramer W. Role of land cover changes for atmospheric CO2 increase and climate change during the last 150 years. Glob. Change Biol, 2004, vol. 10, pp. 1253-1266.
            
          </mixed-citation>
     <mixed-citation xml:lang="en">
            
              Brovkin V., Sitch S., Bloh von W., Claussen M., Bauer E., Cramer W. Role of land cover changes for atmospheric CO2 increase and climate change during the last 150 years. Glob. Change Biol, 2004, vol. 10, pp. 1253-1266.
            
          </mixed-citation>
    </citation-alternatives>
   </ref>
   <ref id="B33">
    <label>33.</label>
    <citation-alternatives>
     <mixed-citation xml:lang="ru">
            
              Stainforth D.A., Aina T., Christensen C. et al. Uncertainty in predictions of the climate response to rising levels of greenhouse gases. Nature, 2005, vol. 433, pp. 403-406.
            
          </mixed-citation>
     <mixed-citation xml:lang="en">
            
              Stainforth D.A., Aina T., Christensen C. et al. Uncertainty in predictions of the climate response to rising levels of greenhouse gases. Nature, 2005, vol. 433, pp. 403-406.
            
          </mixed-citation>
    </citation-alternatives>
   </ref>
   <ref id="B34">
    <label>34.</label>
    <citation-alternatives>
     <mixed-citation xml:lang="ru">
            
              Fitter A.H., Self G.K., Brown T.K., Bogie D.S., Graves J.D., Benham D., Ineson P. Root production and turnover in an upland grassland subjected to artificial soil warming respond to radiation flux and nutrients, not temperature. Oecologia, 1999, vol. 120, pp. 575-581.
            
          </mixed-citation>
     <mixed-citation xml:lang="en">
            
              Fitter A.H., Self G.K., Brown T.K., Bogie D.S., Graves J.D., Benham D., Ineson P. Root production and turnover in an upland grassland subjected to artificial soil warming respond to radiation flux and nutrients, not temperature. Oecologia, 1999, vol. 120, pp. 575-581.
            
          </mixed-citation>
    </citation-alternatives>
   </ref>
   <ref id="B35">
    <label>35.</label>
    <citation-alternatives>
     <mixed-citation xml:lang="ru">
            
              Liski J., Ilvesniemi H., Makela A., Westman C.J. СО2 emissions from soil in response to climatic warming are overestimated - The decomposition of old soil organic matter is tolerant of temperature. Ambio, 1999, vol. 28, pp. 171-174.
            
          </mixed-citation>
     <mixed-citation xml:lang="en">
            
              Liski J., Ilvesniemi H., Makela A., Westman C.J. SO2 emissions from soil in response to climatic warming are overestimated - The decomposition of old soil organic matter is tolerant of temperature. Ambio, 1999, vol. 28, pp. 171-174.
            
          </mixed-citation>
    </citation-alternatives>
   </ref>
   <ref id="B36">
    <label>36.</label>
    <citation-alternatives>
     <mixed-citation xml:lang="ru">
            
              Rochette P., Angers D.A., Flanagan L.B. Maize Residue Decomposition Measurement Using Soil Surface Carbon Dioxide Fluxes and Natural Abundance of Carbon-13. Soil Science Society of America Journal, 1999, vol. 63, pp. 1385-1396.
            
          </mixed-citation>
     <mixed-citation xml:lang="en">
            
              Rochette P., Angers D.A., Flanagan L.B. Maize Residue Decomposition Measurement Using Soil Surface Carbon Dioxide Fluxes and Natural Abundance of Carbon-13. Soil Science Society of America Journal, 1999, vol. 63, pp. 1385-1396.
            
          </mixed-citation>
    </citation-alternatives>
   </ref>
   <ref id="B37">
    <label>37.</label>
    <citation-alternatives>
     <mixed-citation xml:lang="ru">
            
              Risk D., Kellman L., Beltrami H. Carbon dioxide in soil profiles: Production and temperature dependence. Geophysical Research Letters, 2002, vol. 29, no. 6, pp. 111-114.
            
          </mixed-citation>
     <mixed-citation xml:lang="en">
            
              Risk D., Kellman L., Beltrami H. Carbon dioxide in soil profiles: Production and temperature dependence. Geophysical Research Letters, 2002, vol. 29, no. 6, pp. 111-114.
            
          </mixed-citation>
    </citation-alternatives>
   </ref>
   <ref id="B38">
    <label>38.</label>
    <citation-alternatives>
     <mixed-citation xml:lang="ru">
            
              Piao S., Ciais P., Friedlingstein P. et al.Net carbon dioxide losses of northern ecosystems in response to autumn warming. Nature, 2008, vol. 451, pp. 49-52.
            
          </mixed-citation>
     <mixed-citation xml:lang="en">
            
              Piao S., Ciais P., Friedlingstein P. et al.Net carbon dioxide losses of northern ecosystems in response to autumn warming. Nature, 2008, vol. 451, pp. 49-52.
            
          </mixed-citation>
    </citation-alternatives>
   </ref>
   <ref id="B39">
    <label>39.</label>
    <citation-alternatives>
     <mixed-citation xml:lang="ru">
            
              Wang G., Li F., Peng Y. et al. Responses of soil respiration to experimental warming in an alpine steppe on the Tibetan Plateau. Environ. Res. Lett., 2019, vol. 14, p. 10.
            
          </mixed-citation>
     <mixed-citation xml:lang="en">
            
              Wang G., Li F., Peng Y. et al. Responses of soil respiration to experimental warming in an alpine steppe on the Tibetan Plateau. Environ. Res. Lett., 2019, vol. 14, p. 10.
            
          </mixed-citation>
    </citation-alternatives>
   </ref>
   <ref id="B40">
    <label>40.</label>
    <citation-alternatives>
     <mixed-citation xml:lang="ru">
            
              Brovkin V. Climate-vegetation interaction. J. Phys. IV France, 2002, vol. 12, pp. 52-57.
            
          </mixed-citation>
     <mixed-citation xml:lang="en">
            
              Brovkin V. Climate-vegetation interaction. J. Phys. IV France, 2002, vol. 12, pp. 52-57.
            
          </mixed-citation>
    </citation-alternatives>
   </ref>
   <ref id="B41">
    <label>41.</label>
    <citation-alternatives>
     <mixed-citation xml:lang="ru">
            
              Pritchard S.G., Davis M.A., Mitchell R.J., Prior A.S., Boykin D.L., Rogers H.H., Runion G.B. Root dynamics in an artificially constructed regenerating longleaf pine ecosystem are affected by atmospheric CO2 enrichment. Environmental and Experimental Botany, 2001, vol. 46, pp. 35-69.
            
          </mixed-citation>
     <mixed-citation xml:lang="en">
            
              Pritchard S.G., Davis M.A., Mitchell R.J., Prior A.S., Boykin D.L., Rogers H.H., Runion G.B. Root dynamics in an artificially constructed regenerating longleaf pine ecosystem are affected by atmospheric CO2 enrichment. Environmental and Experimental Botany, 2001, vol. 46, pp. 35-69.
            
          </mixed-citation>
    </citation-alternatives>
   </ref>
   <ref id="B42">
    <label>42.</label>
    <citation-alternatives>
     <mixed-citation xml:lang="ru">
            
              Morgan J.A., LeCain D.R., Mosier A.R., Milchunas D.G. Elevated CO2 enhances water relations and productivity and affects gas exchange in C3 and C4 grasses of the Colorado shortgrass steppe. Global Change Biol., 2001, vol. 7, pp. 451-466.
            
          </mixed-citation>
     <mixed-citation xml:lang="en">
            
              Morgan J.A., LeCain D.R., Mosier A.R., Milchunas D.G. Elevated CO2 enhances water relations and productivity and affects gas exchange in C3 and C4 grasses of the Colorado shortgrass steppe. Global Change Biol., 2001, vol. 7, pp. 451-466.
            
          </mixed-citation>
    </citation-alternatives>
   </ref>
   <ref id="B43">
    <label>43.</label>
    <citation-alternatives>
     <mixed-citation xml:lang="ru">
            
              Gifford R.M. Implications of CO2 effects on vegetation for the global carbon budget. In: The global carbon cycle (ed. M. Heimann). Springer-Verlag, Berlin, 1993, pp. 159-199.
            
          </mixed-citation>
     <mixed-citation xml:lang="en">
            
              Gifford R.M. Implications of CO2 effects on vegetation for the global carbon budget. In: The global carbon cycle (ed. M. Heimann). Springer-Verlag, Berlin, 1993, pp. 159-199.
            
          </mixed-citation>
    </citation-alternatives>
   </ref>
  </ref-list>
 </back>
</article>
